The tiny moths in this large family (122 NA spp) are primarily leaf miners
that pupate outside the mine. Three common genera are Lyonetia,
Bucculatrix and Phyllocnistis. Bucculatrixpomifoliella makes a serpentine mine on apple leaves. It
overwinters in a small longitudinally ribbed coccoon on apple twigs. Phyllocnistispopuliella makes a serpentine mine on trembling aspen. Lyonetia
alniella makes a blotch mine on alder. Other species of Lyonetia
occur on Ceanothus and Vitis.